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Monday, June 17, 2019

Arena and Vitale (2018) -- Elevated CO2 Reduces Cold Temperature Stress on Plants

Arena, C. and Vitale, L. 

2018

"Chilling-induced
reduction of
photosynthesis 
is mitigated by 
exposure to 
elevated CO2 
concentration"

Photosynthetica 56: 1259-1267



NOTE:
Multiple research studies 
have shown that elevated levels 
of atmospheric CO2 can alter 
important physiological processes 
to help lessen or fully ameliorate 
the negative impacts of 
high temperature stress 
on plants.

Very little research has been
conducted on the ability of 
higher levels of CO2 to reduce 
the negative impacts 
of cold temperature stress. 

Arena and Vitale (2018) say 
cold temperature stress 
"represents a critical 
environmental stress 
that limits 
plant productivity 
and distribution 
in many regions 
of the world."



SUMMARY:
In response to chilling stress, 
plants in the elevated CO2 treatment 
exhibited higher values of CO2 fixation, 
photochemical activity and 
Rubisco amount than plants 
exposed to ambient CO2. 

Arena and Vitale say the reason 
for this difference was: 

(1) 
Under ambient conditions, 
the "increase of thermal 
dissipation processes 
was not adequate 
to guarantee the PSII 
photoprotection and 
the phososynthetic 
recovery after chilling," 
but 

(2) 
"The elevated CO2 concentration 
in combination with low temperature 
exerted a mitigating effect 
on reduction of Rubisco content 
induced by chilling," 
   which allowed leaves 
"to maintain higher 
photosynthetic rates 
and reduce the adverse effect 
of low temperature on 
Rubisco oxygenase activity, 
upon return to 
prechilling conditions." 

Arena and Vitale conclude 
that their study reveals 
"the positive effect 
of elevated CO2 
in mitigating 
chilling stress." 



DETAILS:
Chilling adversely impacts 
key physiological processes. 

Even a small 
drop in temperature, 
which does not produce 
visible damage to a plant, 
can "induce up to [a] 
50% reduction 
of plant productivity" 
  by altering a plant's 
"water uptake, mineral nutrition, 
photosynthesis, respiration, 
and total metabolism." 

Given the large volume 
of research that demonstrates 
the benefits of elevated levels 
of atmospheric CO2 
on these processes, 
the two Italian researchers 
 set out to examine 
"the role of elevated CO2 
concentrations in mitigating 
chilling-induced photosynthetic
 depression in Beta vulgaris L., 
a winter crop widely cultivated 
in the Mediterranean 
ago-ecosystems."

Arena and Vitale grew chard 
( B. vulgaris cv. Cicla ) 
in a controlled-environment 
at the Department of Biology 
of the University of Naples 
Federico II, Italy, under 
well-watered and fertilized 
conditions. 

Goups of plants were exposed 
to one of four five-hour treatments: 
(1)
 ambient CO2 (360 ppm) 
and ambient temperature (25°C), 

(2)
elevated CO2 (700 ppm) 
and ambient temperature (25°C), 

(3)
 ambient CO2 (360 ppm) 
and low temperature (4°C), 
and

(4) 
elevated CO2 (700 ppm) 
and low temperature (4°C). 

During and following 
the short-term treatment, 
multiple measurements 
were made on key processes 
relating to photosynthesis
to determine if elevated CO2 
was able to reduce the impacts 
of chilling stress.